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SUMMARY
Giant carnivorous dinosaurs such as Tyrannosaurus rex and abelisaurids are characterized by highly reduced forelimbs that stand in contrast to their huge dimensions, massive skulls, and obligate bipedalism. Anothergroup that follows this pattern, yet is still poorly known, is the Carcharodontosauridae: dominant predatorsthat inhabited most continents during the Early Cretaceous and reached their largest sizes in Aptian-Cenomanian times. Despite many discoveries over the last three decades, aspects of their anatomy,especially with regard to the skull, forearm, and feet, remain poorly known. Here we report a new carcharodontosaurid, Meraxes gigas, gen. et sp. nov., based on a specimen recovered from the Upper Cretaceous Huincul Formation of northern Patagonia, Argentina. Phylogenetic analysis places Meraxes among derived Carcharodontosauridae, in a clade with other massive South American species. Meraxes preserves novelanatomical information for derived carcharodontosaurids, including an almost complete forelimb that provides evidence for convergent allometric trends in forelimb reduction among three lineages of large-bodied,megapredatory non-avian theropods, including a remarkable degree of parallelism between the latestdiverging tyrannosaurids and carcharodontosaurids. This trend, coupled with a likely lower bound on forelimb reduction, hypothesized to be about 0.4 forelimb/femur length, combined to produce this short-armedpattern in theropods. The almost complete cranium of Meraxes permits new estimates of skull length in Giganotosaurus, which is among the longest for theropods. Meraxes also provides further evidence thatcarchardontosaurids reached peak diversity shortly before their extinction with high rates of trait evolutionin facial ornamentation possibly linked to a social signaling role. RESULTS: Systematic paleontology: Theropoda Marsh, 1881.Tetanurae Gauthier, 1986. Allosauroidea Marsh, 1878. Carcharodontosauridae Stromer, 1931. Carcharodontosaurinae Stromer, 1931. Giganotosaurini Coria and Currie, 2006. Meraxes gigas gen. et sp. nov.Etymology: Meraxes after a dragon of the Song of Ice and Fire.
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S America was home to countless sauropods so there were carnivorous dinos to feed on them. Meraxes, even though it belongs in a different family than T. rex, shows the same evolutionary trend toward reduced fore lims in these apex predators.